Labour deputy leader Angela Rayner wrote to Simon Case, the head of the civil service, asking him to investigate answers about the affair given by Johnson’s then-press secretary earlier this year. Allegra Stratton, a former BBC journalist, was Johnson’s press secretary from October until last week.

Dominic Cummings, who was Johnson’s key adviser on the Brexit campaign and helped him to win an election in 2019 before an acrimonious split last year, said on Friday that Johnson had wanted donors to pay for the renovation secretly.

Cummings said he had told the prime minister such plans were “unethical, foolish, possibly illegal”.

Asked if Johnson had received a loan from the Conservative Party for the refurbishment, transport minister Grant Shapps told Sky News: “The prime minister has already paid for it.”

In a further potentially damaging allegation, the Daily Mail newspaper on Sunday cited unidentified sources as saying that, in October, shortly after agreeing to a second lockdown, Johnson had told a meeting in Downing Street: “No more f***ing lockdowns — let the bodies pile high in their thousands.”

Asked on Monday whether he had made the remark, Johnson told broadcasters: “No, but again, I think the important thing, I think, that people want us to get on and do as a government is to make sure that the lockdowns work, and they have.” 

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